Přehled
Students are trained in all areas of biochemistry, including bioenergetics, enzymology, molecular biology, protein chemistry and structure, immunochemistry, xenobiochemistry, and metabolic processes. Training involves participation in scientific projects, mainly at the Department of Biochemistry at Charles University and in collaboration with research institutes of the Czech Academy of Sciences and other laboratories. The study concludes with an English language certificate, a state doctoral exam in biochemistry, and a successful dissertation defense with a publication in an international journal. The program is conducted without specialization.
Start of studies in the summer semester (April)
Opening of applications and publication of the dissertation topics: 15. 11. 2025
Application by: 15. 12. 2025
Detailed information about the study programme
Offer of dissertation topics:
- Structural Immunology of Natural Killer Cell Receptor:Ligand Recognition
- Structural mass spectrometry of proteins and protein-nucleic acid complexes
- The structure-functional study of proteins and its complexes – employing of non-canonical aminoacids and protein chemistry combined with mass spectrometry
Profile of the graduate
The graduates of the Ph.D. program in Biochemistry have broad theoretical and practical knowledge of biochemistry and molecular biology. They are prepared for individual scientific project or cooperation in research group. They are able to design and perform complex biochemical experiments and to adapt and apply them relevantly to any chemical and biological topics. The students are able to present their results, critically evaluate them and to discuss them in respect to international scientific context and quality. The in-depth knowledge not only makes them expert in biochemistry and molecular biology with the well-founded scientific orientation in majority of relative topics. The comprehensive understanding enables students to apply and transfer obtained skills during education at University, and also they are able to extract, simplify and share achieved results to broad scientific as well as laic audiences. In addition to biochemical expertise students also are able to develop their scientific, pedagogical and language skills, presentation and rhetorician proficiency, team work and scientific co-operation during the studies. Therefore, the absolvents are able to serve as principal investigators and leader of research groups at scientific institutions as well as specialists at clinical chemistry laboratories, to be employed educational and governmental institutions as well as to participate in specialized applied research, biotechnological, food and pharmaceutical industry or business trade.
Application submission
The application is submitted electronically through the Charles University study information system. The completed application must be submitted electronically in the study information system no later than on 30. 4. 2026 for study programmes starting in the winter semester (October) and no later than on 15. 12. 2025 for study programmes starting in the summer semester (April).
The admission fee is 100 EUR and it is paid for each submitted application.
Required application appendice is the agreement of a supervisor in the format of the following form available here. We recommend reaching out to the supervisor as soon as possible (ideally before 31. 3. 2026 for study programmes starting in the winter semester and before 30. 11. 2025 for study programmes starting in the summer semester), so the application could be send out in time in case the agreement is granted.
Entrance exam
The regular date of entrance exam for all doctoral programmes starting in the winter semester (October) is in June 2026 (to be specified), an alternative date is in June 2026 (to be specified).
The regular date of entrance exam for all doctoral programmes starting in the summer semester (April) is 4. – 5. 2. 2026, an alternative date is 11. – 12. 2. 2026.
The request for an alternative day is submitted and approved via the electronic information system.
The form of the entrance exam is an oral interview, which mainly concerns the issues of the planned dissertation and the candidate’s knowledge within the field of expertise of the given study programme.
Find more information in:
Important links
Applicants with specific needs
If you have a physical handicap, you can apply for a modification of the entrance exam. For more information read the Conditions of the admission procedure for doctoral study programmes.
Departments involved in this programme
Guarantor of the study programme
doc. RNDr. Miroslav Šulc, Ph.D.
miroslav.sulc@natur.cuni.cz
Do you have a question?
Complete information about the admission procedure can be found on the website for applicants. For questions, please contact the admissions contact persons.
